Subspecies:

Acanthodactylus scutellatus hardyi  HAAS, 1957

Types:

Holotype: HUJ Hebrew University of Jerusalem 2682

Terra Typica:

`Hirmas Station` in northern Saudi Arabia.

Taxonomic notes:

Acanthodactylus scutellatus hardyi was described in 1957 by HAAS from northern Saudi Arabia. HARRIS & ARNOLD raised the taxon 2000 without any justification to the status of species: `... previously treated as subspecies are now better regarded as full species (now A hardyi ; for differences from A. scutellatus , see ARNOLD 1986 ...) `, a procedure that was adopted by later authors without criticism. Until a reasoned decision, the taxon will continue to be considered a subspecies here.

WERNER MAYER, 2015

Relevant taxonomic literature:

• Haas, G. (1957) -  Some amphibians and reptiles from Arabia. -  Proceedings of the California Academy of Sciences, 4th ser., 29: 47-86.  

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• Harris, D.J. & Arnold, E.N. (2000) -  Elucidation of the relationships of spiny-footed lizards, Acanthodactylus spp. (Reptilia: Lacertidae) using mitochondrial DNA sequence, with comments on their biogeography and evolution. -  Journal of Zoology, London, 252 (3): 351-362.    

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  Mitochondrial DNA sequences consisting of 645 sites from the 12S rRNA and 16S rRNA genes were used to estimate the phylogeny of 15 of the 32 species of spiny-footed lizards Acanthodactylus. The resultant tree has similarities to that produced from a differentially weighted data set of 32 morphological characters but there are also significant differences. However, combined analysis of molecular and morphological data sets produces the same tree topology as DNA sequence alone. The molecular data confirm that there are distinct eastern and western clades within Acanthodactylus, but place A. boskianus in the former while the A. scutellatus group constitutes a third clade. Species for which only morphological information is available were integrated with the combined tree to give a provisional phylogeny for 31 species. This phylogeny indicates that the ancestor of existing Acanthodactylus probably originated in south-west Asia and that North Africa was invaded by more than one lineage of the genus. It also suggests that soft aeolian sand habitats may have been independently occupied more than once. Molecular data provide independent evidence that the differential weighting of morphological characters in past analyses was appropriate.