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Species: |
| arguta (PALLAS, 1773) |
Subspecies (5): |
| arguta arguta (PALLAS, 1773) arguta darevskii TSARUK, 1986 arguta deserti (GMELIN, 1789) arguta transcaucasica DAREVSKY, 1953 arguta uzbekistanica CHERNOV, 1934 |
Synonyms: |
| Lacerta arguta PALLAS, 1773 Lacerta deserti GMELIN, 1789 Lacerta variabilis PALLAS, 1814 Lacerta leucosticta LICHTENSTEIN, 1823 Lacerta pustulata EICHWALD, 1831 Podarcis irritans MENETRIES, 1832 Lacerta boristhenis DVIGUBSKY, 1832 Eremias variabilis WIEGMANN, 1834 Podarcis deserti EICHWALD, 1841 Eremias arguta GRAY, 1845 Podarcis arguta STRAUCH, 1876 Ommateremias arguta LANTZ, 1928 Podarcis variabilis WAGLER, 1830 |
Common names: |
| Steppe Runner (English)
Steppenrenner (German)
Разноцветная ящурка (Russian) |
Types: |
| arguta: Neotype: ZISP (ZIL) 13205 (designated by Szcerbak 1974:149); Holotype: unlocated fide I. Doronin (pers. comm., Nov. 2018)
darevskii: Holotype: ZISP 20018, Kyrgyzstan, Issyk-Kul Basin, Karakol (formerly Przhevalsk), 1906, Coll. D.D. Pedashenko
transcaucasica: Holotype: ZMMU (also given as ZM MSU) Re 10098, 8.09.1952, Coll. I.S. Darevsky
uzbekistanica: Holotype: ZISP 10093, Uzbekistan, Golodnaya Steppe Station, Central Asian Railway, 05.1903, Coll. G.G. Jacobson |
Terra Typica: |
| arguta: Type locality: between Ural River and Emba River, NW Kazakhstan [see Zhao and Adler, 1993:202 and Bauer and Günther, 1995:56 for additional comments].
darevskii: Type locality: Karakol (formerly Przhevalsk), Issyk-Kul Basin, Kyrgyzstan.
deserti: Type locality: Yaik steppe (between the Volga and the Urals).
transcaucasica: Type locality: Armenia, Basargechar district, environs p. Mets-Mazra.
uzbekistanica: Type locality: Golodnaya Steppe, Uzbekistan. |
Taxonomic notes:
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| Subgenus Eremias sensu stricto |
Relevant taxonomic literature:
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Pallas, P.S. (1773) - Description of Eremias arguta arguta. - In: “Reise durch verschiedene Provinzen des Russischen Reiches. Vol. 2”. - Kaiserlichen Akademie der Wissenschaften, St. Petersburg. 744 pp.  Bauer, A.M. & Günther, R. (1995) - An annotated type catalogue of the lacertids (Reptilia: Lacertidae) in the Zoological Museum, Berlin. - Mitteilungen aus dem Zoologischen Museum in Berlin, 71: 37-62.  × A complete catalogue is provided for the type specimens of lacertid lizards in the herpetological collection of the Zoological Museum, Berlin, as of December, 1993. The collection contains a total of 514 type specimens, representing 63 taxa, of which 42 are currently regarded as valid at the specific or subspecific level. Types representing an additional four taxa appear to have been lost from the collection. The collections are especially rich in African lacertid types and in historically significant specimens collected early in the Nineteenth Century by Pallas, Eversmann, and Hemprich & Ehrenberg. Also present are the type series of many subspecies of Podarcis lilfordi and P. pityusensis described by Eisentraut. Chirikova, M.A. & Kolbinzev, W.G. (2003) - Morphologie und Unterartgliederung des Steppenrenners, Eremas arguta PALLAS, 1773, in Kasachstan. - Salamandra, 39 (1): 49-60.  × Morphology and subspecies of Eremias arguta (PALLAS, 1773), in Kazakhstan.
The external morphology of Eremias arguta in Kazakhstan was examined. Three metric and ten meristic characters as well as the dorsal pattern of 206 individuals from 13 populations of southern, south-eastern and eastern Kazakhstan were analyzed. Sexual dimorphism was found to occur in relative tail length (fem. > males), number of longitudinal gular scales (fem. < males), and number of femoral pores (fem. < males). Clinal variation exists in the number of dorsals, gulars, femoral pores, and scales between femoral pores. Most of populations studied revealed a pattern of large white
spots and transverse dark strips. Some specimens from eastern Kazakhstan are characterized by small white spots and interrupted, somewhat fused dark strips. Most southern populations show elongated white spots and irregular black spots scattered along the dorsum. Subspecies status of the steppe-runners from eastern Kazakhstan is still unclear and needs future clarification. Populations from south-eastern Kazakhstan are closer to E. arguta potanini. Southernmost populations belong to E. a. uzbekistanica. Poyarkov Jr., N.A. & Orlova, V.F. & Chirikova, M.A. (2014) - The mitochondrial phylogeography and intraspecific taxonomy of the Steppe Racerunner, Eremias arguta (Pallas) (Lacertidae: Sauria: Reptilia), reflects biogeographic patterns in Middle Asia. - Zootaxa, 3895 (2): 208-224. × Steppe racerunner, Eremias (Eremias) arguta, is one of the most widespread species of the Asian racerunners (genus Er-emias). Several subspecies were traditionally recognized however, morphological variability is so high that delimitation of these subspecies was always problematic. Here we present a phylogenetic hypothesis for this species based on cyto-chrome b sequences (55 samples from 35 populations, 900 bp partial sequences), infer it biogeography and the revise its subspecific structure. Six major phylogenetic lineages were revealed. The southernmost populations (E. a. uzbekistanica) from Uzbekistan form a clade together with the Issyk-Kul Lake subspecies (E. a. darevskii) based on both molecular and morphological evidence. Within more northern populations, there is a split between populations from Northern Caucasus, Europe and Western Kazakhstan (E. a. deserti) and Central and Eastern Kazakhstan populations (E. a. arguta). Transcau-casian (E. a. transcaucasica) steppe racerunners are grouped with Middle Asian populations. Finally, the easternmost samples, assigned to “E. a. potanini” are nested within the E. a. arguta clade. Populations from the Ili River Valley form a separate lineage sister to the clade joining all other E. arguta lineages and might represent a yet undescribed taxon. Spe-cies distribution in relation to historical biogeography of Middle Asia is discussed.
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